­F

¨å«¬ªº­F¡]2N¡A¥ÀÅé»P¤÷Åé¦U¥b®M¿ò¶Ç½è¡^¡A¥]¬A­F¶b¤Î¨ä¤Wªº¤l¸­¡C­F¶b¥Ñ­FªÞ(Plumule)¡B¤W­F¶b(epicotyl)¡B¤U­F¶b(hypocotyl)¡B©M­F®Ú(radicle)©Ò²Õ¦¨¡C¤U­F¶b»P­F®Úªº¤À¬ÉÂI³q±`¤£©ö°Ï§O¡C¤l¸­ªþµÛ¦b¤U­F¶b¡C¤l¸­ªº¼Æ¥Ø¦]´Óª«ºØÃþ¦Ó²§¡C¤j¦h¼ÆªºÂù¤l¸­´Óª«¬Ò¦³¨â¤ù¤l¸­¡C»r¤l´Óª«¤¤°£conifer ¦³¸û¦hªº¤l¸­¤§¥~¡A¤j¦h¤]¬O¨ã¦³¨â¤ù¤l¸­¡C

¥Ý¥»¬ìªº­F¦b³æ¤l¸­´Óª«·í¤¤¸û¬°¯S®í¡C­F»P­F¨Å³s±µ³B¦³­Ó¯S®í¤Æªº²Õ´¡AºÙ¬°­F½L(scutellum)¡A±`³Q»{¬°¬O¥Ý¯óºØ¤lªº¤l¸­¡C­F½Lªø¦Ó¦ü¬Þ¡A¨ä¤¤¨ã¦³ºûºÞ§ô²Õ´¡C­F½L±µÄ²­F¨Åªº¨º­±¦³¤@¼h¥q¤Àªc§l¦¬ªºªí¥Ö²Ó­M(epithelial cell)¡C

­F½L»P­F³s±µ³BºÙ¬°­F½L¸`(scutellar node)¡C­F¥Ñ¦¹¤À¬°¤W¤U¨â¥b¡C¤W¥b³B¥iºÙ¬°¤W­F¶b¡A¥]¬A¤@¨Ç¸­­ìÅé©M²ô³»¥ÍªøÂI(Shoot apex)¡A¤W­F¶b¥Ñ¤@¤¤ªÅª¬ª«©Ò¥]³ò¡A§Y¬OÀT¸­(coleoptile)¡A³o¤@¨Ç¾¹©x¦XºÙ¬°acrospire¡C¥¿¹ïµÛ­F½L¸`¡A¬Û¹ï©ó­F½Lªº¥t¤@°¼¡A¥i¯àªø¥X¤@¤£¨ãºûºÞ§ôªº¤p¬ð°_¡AºÙ¬°epiblast¡]¦p¤p³Á¡B¿P³Á¦³¤§¡A¦ý¥É¸¾¶Á«h¨S¦³¡^¡A¦³¾ÇªÌ»{¬°¬O¥t¤@¥b¤l¸­ªº²ª¸ñ¡A¥i§@¬°³æ¤l¸­¥ÑÂù¤l¸­¶i¤Æ¦Ó¨ÓªºªºÃÒ¾Ú¡CÀT¸­°ò³¡»P­F½L¤§¶¡¡Aµu¤pªº²Õ´ºÙ¬°¤¤­F¶b(mesocotyl)¡B­F¶b¦b­F½L¸`¤§¤UºÙ¬°­F®Ú(radicle)¡B­F®Ú¤§¥~¥]³ò¤@¼h²Õ´¡A§Y¬OÀT®Ú(coleorhiza)¡C

¥Ý¯ó­F¤§ºc³y¦b¹L¥h150 ¦~¥H¨Ó¡A¤@ª½¬°¾ÇªÌª§½×¤§ÂI¡C­F½L¤@¯ë»{¬°¬O¤l¸­¡A¦Óepiblast «h¬O¥t¤@¤ù¤w°h¤Æªº¤l¸­¡AÀT¸­«h¬O¥t¤@ºØ¸­ªºÅܧΡC¦ÓÀT®Ú¬°primary root¨üÀ£§íÅܧΦӦ¨¡C¦ý¥ç¦³¤Ö¼Æ¾ÇªÌ±NÀT¸­»{¬°¬O­F½L¤§ÀT¡BµÅªÞ¡A©Î¬O¬Æ¦Ü»{¬°ÀT¸­§Y¬O¤l¸­¡C¦Óepiblast«h¬OÀT¸­¤§¥~¥Íª«¡C¦Ü©ó¤¤­F¶bªº°_·½¡Aª§¯É¥ç¦h¡A©Î»{¬°¬O¤@¸`¶¡¡B©Î»{¬°¬O¸`¡A©Î¥H¬°¬O¤U­F¶b»P­F½L±µµÛ¿Ä¦X¦Ó¦¨¡A©ÎªÌ¬°¤l¸­¤§¤@³¡¥÷¤£µ¥¡C

­F¤§ºc³y¦b¥Ý¥»¬ì¤§¤º¨ã¦³¤ÀÃþ¨t²Î¤Wªºª[²§¡ARecder±N150 ÄÝ300 ºØ¥Ý¯óª«¨Ì¤U¦C¥|­Ó¯S©Ê¤À¦¨ 6¸s¡A1¡G­F½L»P­ìÅéºûºÞ§ô¤Àª[¤§¦ì¸m¡A2¡Gepiblast¤§¦s¦b»P§_¡A3¡G­F½L¤§©³³¡»PÀT®Ú³s±µ»P§_¡A4¡G­F¸­¾î¤Á­±ªº¯S¼x¡C

Epiblast¦b¤ÀÃþ¤Wªº¥\¯à¬Æ¦h¡A¨Ò¦pBrakworth¡]1982¡^µo²{epiblast¤§¯S©Ê»P­F¤§ªø¼e¤ñ¥i¥Î¨Ó°Ï¤ÀStipa©MOryzopsis¡BTateoka¡]1964¡^§Q¥Îepiblastªº¯S©Ê»Pauricle¤§¦³µL±NOryza ¡]½_ÄÝ¡^¤À¦¨¤T­Ó¸s¡A¦ÓBuam¡]1986¡B1978¡^«h§Q¥Îepiblastªº§ÎºA±NAvena fatua»PA. sativa¡FHordeum brachyantherum»PH. secalinum°Ï¤À¡C

¡@

µo¨|¤£¥þ­F

¤j¦h¼ÆªººØ¤l¡A¬Ò¦³µo¨|§¹¥þªº­F¡A¥]§t­F¶b»P¤l¸­¡B­F¶b¨âºÝ¦U¦³³»ºÝ¤À¥Í²Õ´¡A¤l¸­¸`¤§¤W¬°¤W­F¶b¡V­FªÞ¡F¤l¸­¸`¤§¤U¬°¤U­F¶b¡V­F®Ú¡C¦ý³¡¥÷´Óª«ªº­F«hµo¨|¤£¥þ¡A­F¤p¦Ó¤£¨ã¦³¤l¸­©Î³»ºÝ¤À¥Í²Õ´¡A¨¥¨Ç­F§Y¨Ï¦bºØ¤l¦¨¼ô´²¼½®É¡A¤´µM¥¼¶i¤@¨Bµo¨|¡A³oºØºØ¤l¦b³æ¤l¸­¤ÎÂù¤l¸­´Óª«¬Ò¦³¤§¡C

³Ì²³æªº­F­nºâMenotropa uniflora¡A¨ä­F¡]¥u¯à»¡¬O­ì­F¡^¥u§t¨â­Ó²Ó­M¡A¨ä¥L´Óª«¥ç¦³¥|¡B¤»¡B¤Q©Î¼Æ¤Q­Ó²Ó­Mªº­ì­FªÌ¡C³oÃþ´Óª«¦h¨£©ó¤£¥þ­F(¦pRanunculaceae¡BPapaveraceae)¡B©³½L­F(¦pSaururaceae¡BXyridaceae)¡BÅÖ«¬­F(¦pBurmanniaceae¡BOrchidaceae) ¤Î·L«¬­F(¦pGentianceae)¡C³o¨Ç´Óª«·í¤¤¦³³¡¥÷¬°saprophytes(¦pGeuitiancea ¡BPyrolaceae¡BBurmanniaceae)©Îparasites(¦pBalonophoracae¡BCyomoriaceae)¡A¦ý¥ç¦³autotrophsªÌ¡C

³o¨Ç­FÁöµMºc³y·¥¬°³æ¯Â¡AµM¦Ó²Ó­M¤ºªº¦¨¤À¤w¸g¤À¤Æ¡A¨Ò¦p¸U¥NÄõªº­F¶È¦³¨â­Ó²Ó­M¡A³»ºÝ²Ó¡]ca¡^Åé¿n¤p¡A¦ý§t¦³°ª¿@«×ªºPNA ©M³J¥Õ½è(A$S)¡C¦Ó¬YºØ¹Å¼wÄR¨ÈÄõ(Cattleya aurantiaca) ªº§Z¶ê§Î­F¡A¨ä³»ºÝ³¡¦ìªº²Ó­M³q±`¸û¤p¡Aª½®|¬ù¬°8-10ƒÝm¡A¦Ó°ò³¡²Ó­M«h¸û¤j¡C

¡@

polymbryo ¦h­F

³q±`¤@²ÉºØ¤l±`§t¦³¤@­Ó­F¡A¦bµoªÞ¤§«á¡Aªø¥X¤@­Ó¥®­]¡CµM¦Ó¦³¨Ç´Óª«±`¥iµo²{¤@²ÉºØ¤l¥i¥Hªø¥X¤G®è¥®­]¡C³o¨ÇºØ¤l°£¤F§t¦³¤@­Ó¨üºë¦Ó¨Óªº­F¤§¥~¡A©|¨ã¦³¨ä¥L¥ÀÅé²Ó­M¤À¤Æ¦Ó¦¨ªº­F¡C

¦h­Fªº°_·½¬ÆÂø¡A¥i¥H¥Ñ¯]¤ß²Ó­M¡B¯]³Q²Ó­Mª½±µ¤À¤Æ¦Ó¨ÓªºµL©Ê­F¡A¤]¥i¯à¥Ñ¦hºë§Zµo¨|¤§­Fªº³»ºÝ²Ó­M©ÎSuspensorµo¨|¦Ó¦¨¡C¦¹¥~§U²Ó­M¤»¥i¯à¸g¨üºë§@¥Î¦Ó²£¥Í¦h­F¡C¨ä¤¤¥H¥Ñ¯]¤ß²Ó­Mµo¨|¦Ó¦¨ªº¦h­F³Ì¬°´¶¹M¡C¬ù¦³16¬ìªº´Óª«±`¥iµo²{¤§¤×¨ä¥HÂcªGAnacardiaceae »P¬a¾ï(ªå­»¬ì)³Ì¬°¤H¼ôª¾¡C

°£¤F¬c¤l¥H¥~¡A¤j³¡¥÷¬a¾ïÃþºØ¤l¬Ò¦³¤l­F²{¶H¡C¥HCitrus microcarpa¬°¨Ò¡A¦b¨üºë¤§«á¡A¯]¤Õªþªñªº¯]¤ß²Õ´§t¦³­Y¤z¨ã¦³¿@²Ó­M½è¡A¤j®Öªº²Ó­M¡C³o¨Ç²Ó­M¸g¤Àµõ¦Ó§Î¦¨¦ü­F­ÓÅé¬ð¤J­FÅn¤§¤º¡C³o¨Ç­ÓÅé¦b­FÅn¤º§l¦¬¾i¤Àµo¨|¦Ó¦¨§¹¥þ­F¡A³o¨ÇµL©Êªº¯]¤ß­F¨ã¦³Suspensor ¦Ó»P¨üºë­F¬Û¦ü¤£©ö°Ï¤À¡C

¥Ñ¬a®Ü¤@¯ë¥HÀç¾i´Ó±ø¦æµL©ÊÁc´Þ¡A¦ý¦¸¼Æ¹L¦h«h´Ó±ø³vº¥¦HÅÜ¡C¦¹®É­Y§Q¥Î¤ß­F¨Ó¶i¦æ§ó·s«h¥i¥H±o¨ì°·§§¦Ó»P¥À¥»¿ò¶Ç²Õ¦¨µL²§ªº´Ó®è¡A¦¹¥~µL©Ê­F¦¨ªøªº´Ó®è¡A¥Ñ©ó¿ò¶Ç²Õ¦¨¤@­P¡A¦]¦¹¥i¥H¥Î¨Ó°µ¬°¹êÅç§÷®Æ¡A¨ä¥D®Ú¨t²Î¤S¤Q¤Àµo¹F¡A¥i¥H´£¨Ñ¬°Àu¨}ªº¯z¤ì¡A¦A²ÉºØ¤l¥i¥H²£¥ÍªºµL©Ê­F¼Æ¥Ø¤£¤@¨Ò¦pCitrus microcarpa¤@²ÉºØ¤l¥i¥H§Î¦¨21­Ó¥®­]¡A¦ÓC. unshiu«h¦h¥i¹F40²É¡C

¥Ñ©ó¬a®ÜµL©Ê­F»P¦³©Ê­F¤£©ö°Ï¤À¡A¦]¦¹¥²»Ý­nµ¥¨ì´Ó®è¬Û·í¦¨¼ô«á¤~©ö©ó¿ë§O¡C¦³¤H§Q¥ÎPoncirus trifoliata¤T½Æ¸­ªº¯S©Ê¨Ó°µ¿ò¶Ç°O¸¹¡A¨Ï¥À®è»PPoncirus trifoliata½Æ¥æ¡A±N¨Ó²£¥Íªº¦h­F¥®­]¤¤¡A­Y¨ã¦³¤T½Æ¸­ªÌ¡A«h¥²¬O¦³©Ê­F¡C¥Ñ©óP. trifoliata¤T½Æ¸­¬O¥Ñ¤@¹ï°ò¦]©Ò±±¨î¡A¦]¦¹¥²¶·¬°¦P½èµ²¦XÅé¤~¯à±oÃÒ§PÂ_µL»~¡C¦¹¥~¤]¥i¥H¥Î¹qªa¤ÀªR¨ä»Ã¯À©Î³J¥Õ½è¡A¥H¸ê¶i¦æµL©Ê­F¤§§PÂ_¡C

¨~ªG¥ç±`¥X²{¦h­FºØ¤l¡A³¡¥÷«~ºØ¤@ºØ¤l¤ºµL©Ê­F°ª¹F50­Ó¡A¦Óµ²¦X­F«h±`°h¤Æ¡C¯]¤ß­F°¸¦Ó¥X²{¤T¤l¸­¡A¬Æ¦Ü©ó¦h­Ó­F³¡¿Ä¦X­±§Î¦¨¤@­Ó®Ú¨t¦Ó¼Æ­Ó²ô³¡ªº¥®­]¡C¨ã§t­F¤§«~ºØ¤@¯ëÄÝ©óªF«n¨È¨t²Î¡A¦p¤gÂcªG¦Ó¦L«×¨t²Î¦p·R¤å¡B®ü¹yªÌ¡A¨S¦³¦h­F¤§ºØ¤l¡C¨ä¥L¦pMyrtaceae(¦pEugenia spp.)¥P¤H´x¬ì(¦pPuntia aillenii)¡B¤ÎÄõ¬ì(¦pZeuxine sclcator)¥ç¬Ò¦³¯]¤ß­Fªº³ø¾É¡C

¡@

Chimeral embryos Æ^­F

Baattaglia¡]1945¡^¦b¬ã¨sRudbeckia laciniata®Éµo²{ªá¯»ºë®Ö¶i¤J§Z²Ó­M¤§«á¡A¤£»P§Z®Ö¿Ä¦X¡C²Ó­M¤Àµõ®É¡A¤G®Ö¦U¦Û¶i¦æ¤Àµõ¡A¾É­P¨ä­F­Ý¨ã¦³¤÷·½¤Î¥À·½²Õ´¡CBaattaglia±N³oºØ­F©R¦W¬°semigamy¡A²{¦b«h¦³¤H±N¤§©ö¦W¬°hemigamy¡CHemigamyªº²{¶H¥X²{¦b¤Ö¼Æ´Óª«¡A¦p´Ö(Gossypium barbadense)ÀA¸ª¬ìMalvaceae¡^¡BRudbeekia spp(Asteraceae)¡BCoperia spp.(¥Û»[¬ìAmaryllidaceae)¡B¤ÎZeplybnthes spp.µ¥¡C

¥Ñ©ó²Ó­M¤Î®Ö¤Àµõªº®É²§¡A¦]¦¹Æ^­Fªº²Ó­M¨ã¦³¦UºØ¥i¯àªº¿ò¶Ç²Õ´¦¨¤À¡A©Î¬°diploid ¶¯®Ö¡A©Î¤ldiploid »Û®Ö¡A¤]¥i¯à¦¨¬°­Ý¨ã¶¯»Û¯S©Êªºhexaploid ¡CÆ^­Mªº¯S©ÊÁöµM¥X²{©ó­FµoªÞ¦­´Á¡A¦ý¨ä¯S©Ê¥i¥H¹ï¶È¥X²{©ó´Ó®è§Î¦¨Æ^´O´Óª«¡A¨Ã¥B¥i¯à¿ò¶Ç¨ì¤U¥N¡C¨Ò¦pPima´Ö¡G±Ndoubled haploidªº«~ºØ¡]§t²`ºñ¦â¸­¤ù»Pªo¸¢Åé¡^»P¥t¤@«~ºØ¡]¸­²Lºñ¦â¡B¤£§tªo¸¢Åé¡^Âø¥æ«á¡A¨ä«á¥N³¡¥÷§e²`ºñ¦â¡A³¡¥÷§e²Lºñ¦â¡A¦³¨Ç¦a¤è¨ãªo¸¢Å馳¨Ç¦a¤è«h§_¡C

¨Ó¦Û¨üºë­Fªº¦h­F¤]¦³¤£¦Pªº°_·½¡C¦bTulioa gesneriana ¡ACocos nucifera¡APrimalus auricula¡ACorydalis cava ¬Ò¬O¥Ñ¨üºë­F³»ºÝ²Ó­M¤Àµõ¦Ó¦¨¦h­F¡C¦ÓCymbidium bicolor«h¥Ñ±µ¦X¤lª½±µ¤Àµõ¦¨¦h­F¡CSarkar and Ce¡]1966¡^¦Û49903­Ó¥É¸¾¶Á·í¤¤µo²{¨ì32­ÓÂù­FºØ¤l¡A¥L­Ì»{¬°¨ä¨Ó·½³¡¥÷¥i¯à¬O­F­ì¤Àµõ¦Ó¦¨¡A©ÎªÌ¬O­FÅn¤º¤G­Ó²Ó­M¸g¨üºë¦Ó§Î¦¨¡C

°£¤F³oºØdiploid-diploid twin¤§¥~¡A©|¦³15­Ó¬Ohaploid¡A¤G­Ó¬Odiploid ¡A¥Ý¥»¬ìÂù­F²{¤µ¤£¥u­­©ó¥É¸¾¶Á¡C§@ªÌ¦b¤j¶q¶i¦æ¤ôºØ¤lµoªÞ¹êÅ礤¡A¥ç°¸¦Óµo²{Âù­F¤ô½_¡CAcanltaceae¤¤±`¥iµo²{¨ì¤@¨Ç¦h­F¡A³£¬O¥Ñsuspensorµo¨|¦Ó¨Óªº¡A¦pDipteracanthus patrlus, Acanthus tilicifolius, Sericocalyx scaber¡C³o¨Ç¥Ñ¨üºë§Z­F©ÎSuspeusor °_·½ªº¦h­F¡A¬Ò§t¦³diploid¬V¦âÅé¥Ñ©ó¦h­F¤¤¡A­¿¼Æ¦³haploid ¡Adiploid¡Atriploid¡Apolyploid µ¥¡C¦]¦¹¨ã¦³¹A·~¤Wªº¥Î³~¡C


¡@